Flightless beetles and stingless bees: phoresy of scotocryptine beetles (Leiodidae) on their meliponine (Apidae)v. 55No. 1125–135
Roubik, David Ward and Wheeler, Quentin D. 1982. "Flightless beetles and stingless bees: phoresy of scotocryptine beetles (Leiodidae) on their meliponine (Apidae)." Journal of the Kansas Entomological Society, 55, (1) 125–135.