Phylogeny and zoogeography of the planktonic copepod genus (calanoida: Tortanidae), with establishment of a new subgenus and descriptions of two new speciesv. 18No. 4774–807
Ohtsuka, S. and Reid, J. W. 1998. "Phylogeny and zoogeography of the planktonic copepod genus Tortanus (calanoida: Tortanidae), with establishment of a new subgenus and descriptions of two new species." Journal of Crustacean Biology, 18, (4) 774–807.