Phylogeny and putative hybridization in the subtribe Paranepheliinae (Liabeae, Asteraceae), implications for classification, biogeography, and Andean orogenyv. 46Wiley Publishing Asia375–390
Soejima, A., Wen, Jun, Zapata, M., and Dillon, M. O. 2008. "Phylogeny and putative hybridization in the subtribe Paranepheliinae (Liabeae, Asteraceae), implications for classification, biogeography, and Andean orogeny." Journal of Systematics and Evolution, 46 375–390.