Systematics of the pentadactylus species group of the frog genus Leptodactylus (Amphibia, LeptodactylidaeDOI: info:10.5479/si.00810282.301Smithsonian Institution Press43
Heyer, W. Ronald. 1979. Systematics of the pentadactylus species group of the frog genus Leptodactylus (Amphibia, Leptodactylidae. Washington: Smithsonian Institution Press. In Smithsonian Contributions to Zoology, 301. https://doi.org/10.5479/si.00810282.301.