Evolution, Biogeography, and Description of a New Species of Fruit-eating Bat, Genus Leach (1821), from Panamáv. 59No. 5257–273
Kalko, Elisabeth Klara Viktoria and Handley, Charles O., Jr. 1994. "Evolution, Biogeography, and Description of a New Species of Fruit-eating Bat, Genus Artibeus Leach (1821), from Panamá." Zeitschrift Für Saügetierkunde, 59, (5) 257–273.